Until Now
BiologyGeology

Footprints before feet

Tracks in a Polish quarry show four-legged animals walking 18 million years before any body fossil.

When
Known to
± 3 million years
Era
Devonian, Paleozoic

Trackways in 395-million-year-old marine tidal-flat carbonates in the Holy Cross Mountains preserve alternating left-right prints with distinct digits — unambiguously made by a tetrapod. They predate the oldest tetrapod skeletons by roughly eighteen million years, and Tiktaalik, the celebrated fish-tetrapod intermediate, by twenty. The find forced a substantial rewrite of when and where vertebrates first walked.

Just before

Just after